Ecology Terms Starting With H
Ecology Glossary: H
Jump to Ecology Term
Habitat Fragmentation
/ HAB-ih-tat frag-men-TAY-shun / · Latin habitare (to inhabit) + fragmentum (broken piece) + -ation
Habitat fragmentation is the process by which a large, continuous area of habitat is divided into smaller, isolated patches separated by inhospitable terrain, reducing the total area available to species and limiting movement between patches.
Roads, agricultural fields, and urban development subdivide once-continuous habitat into isolated patches that may be too small to support viable populations of large-bodied or wide-ranging species. Fragmentation increases edge habitat and perimeter-to-area ratios, exposing interior specialists to harsh microclimate conditions, predation, and invasive species. Gene flow between isolated populations ceases, causing genetic drift and inbreeding that reduce fitness and adaptability even when patches are separated by short distances.
A forest patch must typically exceed 500 to 1,000 hectares to maintain viable populations of many birds and large mammals that require interior habitat conditions. Corridors connecting fragmented patches through continuous or semi-continuous vegetation allow animal movement and genetic exchange between isolated subpopulations.
The Biological Dynamics of Forest Fragments Project, established in the Brazilian Amazon in 1979, is the longest-running fragmentation experiment in the world. Researchers found that forest patches smaller than 100 hectares lost more than half their bird species within 15 years of isolation, with interior specialists disappearing first.
Small isolated habitat patches are safe refuges as long as some habitat remains. Patches below a critical size threshold may support a species temporarily but lose it within years or decades as small populations succumb to inbreeding, demographic accidents, and the absence of recolonization from neighboring areas.
Roads through a forest fragment habitat for amphibians such as spotted salamanders (Ambystoma maculatum), which must cross paved surfaces to reach breeding ponds each spring. Road mortality during these migrations can remove 30 to 50 percent of a local population in a single season, and populations on the road-isolated side of a highway show measurably lower genetic diversity than those with unobstructed access to surrounding forest.
Herbivory
/ HUR-bih-vor-ee / · From Latin herba meaning grass or herb, and vorare meaning to devour
Herbivory is the consumption of plant or algal material by an animal, representing a primary trophic interaction that transfers energy from producers to higher trophic levels.
Herbivory drives ecosystem dynamics by transferring energy from primary producers to higher trophic levels, with herbivores typically consuming only 10 to 20 percent of available plant biomass in most terrestrial ecosystems. Plant defenses against herbivory include physical structures like thorns and silica-reinforced leaf edges, as well as chemical compounds such as tannins, alkaloids, and glucosinolates that reduce digestibility or deter feeding. Different herbivores specialize on multiple plant parts: browsers consume leaves and shoots, grazers eat grasses, and granivores feed on seeds.
Large-scale herbivory can dramatically alter vegetation structure, as seen when white-tailed deer (Odocoileus virginianus) populations reach 30 animals per square kilometer, causing measurable declines in forest regeneration. African savannas demonstrate intensive herbivory where ungulates like zebras (Equus quagga) and wildebeest consume up to 66 percent of annual grass production during seasonal migrations.
Some plants benefit from moderate herbivory. Grasses in particular can grow more vigorously after grazing because removal of older tissue stimulates new growth and increases light reaching the base of the plant; certain prairie ecosystems evolved with sustained grazing pressure from bison, and removing these herbivores can reduce plant diversity over time.
All herbivores damage the plants they feed on. Moderate grazing by insects and mammals can stimulate plant growth, increase plant diversity, and in some cases improve seed dispersal, so the net effect of herbivory on a plant population depends heavily on the intensity and timing of feeding.
The monarch butterfly caterpillar (Danaus plexippus) feeds exclusively on milkweed plants (Asclepias spp.) across North America. Each larva sequesters cardiac glycosides from milkweed tissues, accumulating concentrations toxic enough to cause vomiting in blue jays that attempt to eat them, converting the plant's chemical defenses into the caterpillar's own protection.
Heterotroph
/ HET-er-oh-trohf / · Greek heteros (other) + trophe (nourishment)
Heterotroph is an organism that obtains energy and carbon by consuming organic molecules produced by other organisms, rather than synthesizing them from inorganic sources.
Heterotrophs obtain energy by consuming organic molecules synthesized by autotrophic producers, breaking down complex molecules through cellular respiration to release usable energy. Herbivores such as rabbits consume plant tissues directly, while carnivores such as eagles obtain energy by consuming other animals. Decomposers such as bacteria and fungi secrete enzymes into their substrate, breaking down dead organic matter externally before absorbing nutrient-rich solutions.
Parasitic heterotrophs such as tapeworms absorb pre-digested nutrients across their body surfaces while living inside host organisms. No heterotroph can synthesize organic molecules from inorganic materials alone, which means every heterotrophic organism ultimately depends on autotrophs to fix carbon into usable form.
Some heterotrophs can survive on extraordinarily dilute energy sources: the deep-sea bacterium Candidatus Desulforudis audaxviator lives more than 2.8 kilometers underground in South African gold mines, obtaining energy entirely from hydrogen and sulfate produced by the radioactive decay of uranium in surrounding rock, with no connection to surface photosynthesis.
Mycology →Heterotroph and carnivore mean the same thing. All herbivores, omnivores, decomposers, and parasites are also heterotrophs because they consume organic matter originally produced by other organisms.
A mushroom is a heterotroph that absorbs nutrients from dead organic matter in soil or wood. Its mycelium releases enzymes that break down cellulose and lignin externally, and a single mycelial network can extend across hundreds of square meters while decomposing material that no animal could digest directly.
Are Enzymes Proteins? →Humus
/ HYOO-mus / · From Latin humus meaning earth or soil
Humus is the dark, chemically stable organic component of soil produced when microbial decomposition of plant and animal matter reaches a stage where the remaining compounds resist further rapid breakdown.
Humus represents the end product of decomposition, containing complex organic compounds that can persist in soil for hundreds to thousands of years. Comprising 2 to 10 percent of soil composition in temperate forests, humus improves soil structure by binding mineral particles into aggregates that enhance water retention and aeration. Formation of humus depends on climate, vegetation type, and microbial activity: cool temperate forests produce mor humus that accumulates in distinct layers, while tropical soils often contain minimal humus because warm, moist conditions accelerate decomposition faster than organic matter accumulates.
Humus holds nutrients like nitrogen and sulfur in organic forms, releasing them gradually through mineralization, and its dark coloration comes from humic acids and fulvic acids, complex polymers that also chelate metal ions and buffer soil pH. Carbon stored in humus represents a significant long-term reservoir, with some humic compounds persisting for over 1,000 years as measured by radiocarbon dating.
Humus can hold water up to 90 percent of its own weight, making it far more absorbent than clay or sand. A single gram of humus-rich soil can contain over one billion bacteria, sustaining microbial communities that drive nutrient cycling across entire ecosystems.
Humus and compost are the same material. Compost is partially decomposed organic matter that continues to break down relatively quickly after application to soil, while humus is the chemically stable final product that changes extremely slowly over decades or centuries.
The thick humus layers of boreal forests in Finland and Russia can exceed 20 centimeters in depth, forming over centuries under cool, moist conditions. These organic horizons support distinctive plant communities including Sphagnum mosses, reindeer lichen (Cladonia rangiferina), and ericaceous shrubs such as crowberry (Empetrum nigrum), all adapted to the acidic, nutrient-poor conditions that slow decomposition creates.
