Ecology Terms Starting With R
Ecology Glossary: R
R-Selected Species
/ AR sih-LEK-ted SPEE-sheez / · Latin ratio (rate) + Latin selectio (choice)
R-selected species produce many offspring quickly, investing little energy per individual offspring, and typically experience high juvenile mortality rates.
R-selected species often generate hundreds to millions of offspring per reproductive event, with juvenile mortality reaching 99 to 99.9 percent in many insect and fish populations. Sexual maturity arrives early, sometimes within days to weeks, so populations can double rapidly when resources are temporarily abundant. Small body size and lifespans under one year characterize most r-selected organisms, from annual plants to mayflies.
Unpredictable or frequently disturbed environments favor this strategy because individual survival is unreliable, and population persistence depends on numerical output rather than parental investment.
The common housefly (Musca domestica) can complete its entire life cycle from egg to reproductive adult in as little as 7 to 10 days at warm temperatures, allowing dozens of overlapping generations within a single summer.
R-selected species are evolutionary failures because most of their offspring die. High offspring mortality is precisely what this strategy anticipates; the sheer number of offspring produced ensures that at least some survive to reproduce even when conditions are harsh.
Dandelions release hundreds of wind-dispersed seeds per plant each season, most of which land in unsuitable locations and fail to germinate. Those that land on disturbed soil can establish and flower within weeks, allowing rapid colonization of bare ground before competitors arrive.
Realized Niche
/ REE-uh-lyzd NITCH / · Latin realizare (to make real) + French niche (recess)
Realized Niche is the actual range of environmental conditions and resources a species uses when competition, predation, parasitism, and other biotic interactions are present.
Without competitors or predators, a species can occupy its full fundamental niche, the complete set of conditions it can physiologically tolerate. Biotic interactions compress this potential into a smaller realized niche by excluding the species from portions of its habitat or resource spectrum. The American barnacle (Chthamalus stellatus) can survive across a broad intertidal zone, but competition from the larger barnacle Semibalanus balanoides pushes it upward into the high intertidal, where Semibalanus cannot tolerate desiccation.
Joseph Connell’s classic 1961 experiments on Scottish rocky shores demonstrated this compression by removing Semibalanus and observing Chthamalus expand downward into its full fundamental niche.
Some species have their realized niches shaped not by competitors but by mutualists. The Clark's nutcracker (Nucifrax columbiana) depends so heavily on whitebark pine seeds that its realized niche tracks the distribution of that single tree species across high-elevation western mountains.
A species occupies every habitat it can physically survive in. Predators, competitors, and parasites routinely exclude species from portions of their physiological range, so the realized niche is often much smaller than the fundamental niche.
Barnacles of the genus Chthamalus on rocky shores in the British Isles can physiologically tolerate the full intertidal zone from the splash zone down to the low-tide mark. Competition from Semibalanus balanoides, which grows faster and smothers or undercuts Chthamalus, restricts Chthamalus to the upper 20 to 30 centimeters of the intertidal in areas where both species co-occur.
Reforestation
/ ree-for-es-TAY-shun / · Latin re-, again; forestis, outside (referring to forest); -ation, process
Reforestation is the deliberate or natural reestablishment of tree cover on land that was previously forested but lost its trees through logging, fire, agriculture, or other disturbance.
Reforestation proceeds either through active planting of seedlings or through passive natural regeneration, in which surviving seeds, root systems, and soil seed banks drive recovery without human intervention. Species selection strongly influences outcomes; planting fast-growing exotic monocultures can suppress native understory plants and reduce wildlife habitat value compared with diverse native plantings. Restored forests in the Brazilian Atlantic Forest, one of the world’s most threatened biomes, can recover roughly 80 percent of native bird species within 40 years when planted with a mixture of native tree species.
Soil stabilization, carbon sequestration, and watershed protection all improve as canopy cover and root networks develop over decades.
Costa Rica reversed decades of deforestation between 1985 and 2015, increasing forest cover from roughly 21 percent to over 52 percent of national land area, largely through payments-for-ecosystem-services programs that compensated landowners for allowing natural regeneration.
Types of Trees →Planting any trees anywhere counts as successful reforestation. Species composition, genetic diversity of planting stock, soil preparation, and long-term management determine whether a planted stand develops into a functioning forest ecosystem or remains a low-diversity tree plantation.
Reforestation of degraded pasture in Costa Rica using native canopy species such as guanacaste (Enterolobium cyclocarpum) can attract seed-dispersing birds and bats within two to three years. These animals deposit seeds of additional native species beneath the planted trees, accelerating the recovery of understory diversity beyond what the original planting achieved.
Refugia
/ reh-FYOO-jee-uh / · Latin refugium (place of retreat) + plural -ia
Refugia are localized areas where populations of species persist through periods of regional environmental change that render surrounding habitats unsuitable.
These sheltered areas maintain temperature, moisture, or resource conditions that differ enough from the surrounding regional terrain to buffer resident populations against drought, glaciation, fire, or rapid climate shifts. Mountain valleys, north-facing slopes, deep springs, and cave systems can all function as refugia by preserving microclimatic stability. During the last glacial maximum approximately 20,000 years ago, temperate tree species survived in refugia along the southern margins of European and North American ice sheets, then recolonized northward as glaciers retreated.
Modern climate change is shrinking many existing refugia while creating new ones at higher elevations and latitudes, reshaping where species can persist.
Genetic studies of European brown bears (Ursus arctos) show that nearly all modern European populations descend from a single refugial population that survived the last ice age in the Iberian Peninsula, demonstrating how a single small refugium can be the source of an entire continental recolonization.
Refugia are always large protected parks or nature reserves. A refugium can be a single spring, a shaded rock outcrop, or a small valley a few hectares in area, as long as local conditions remain suitable when the surrounding region becomes inhospitable.
During past glaciations, populations of southern Appalachian brook trout (Salvelinus fontinalis) persisted in cold, spring-fed headwater streams while surrounding lowland waters became too warm. These stream refugia, some less than a few kilometers long, maintained genetically distinct trout populations that later recolonized connected stream networks as temperatures dropped again.
Relict
/ REL-ikt / · Latin relictus (left behind)
Relict is a species or population that persists in a restricted area today after once occupying a much broader geographic range, typically because environmental or climatic changes eliminated it from most of its former distribution.
Relict species are living survivors of lineages that were far more widespread in the past, and their current distributions often map onto refugia where historical conditions have been preserved. The dawn redwood (Metasequoia glyptostroboides) was known only from Cretaceous and Eocene fossils spanning North America and Eurasia until living trees were discovered in a remote valley in Sichuan, China, in 1944. Populations of the horseshoe crab (Limulus polyphemus) along the Atlantic coast of North America represent a morphologically conservative lineage that has changed little in roughly 450 million years, persisting while related groups disappeared.
Studying relict distributions gives ecologists and paleontologists direct evidence of how past climates and habitats differed from present conditions.
The Wollemi pine (Wollemia nobilis), a relict conifer discovered in a sandstone canyon in New South Wales, Australia, in 1994, was previously known only from 90-million-year-old fossils and was assumed extinct. Fewer than 100 wild trees survive in that single canyon.
Relict means extinct. A relict organism is still alive; what has contracted is its range or the broader group to which it belongs, not the organism itself.
The coelacanth (Latimeria chalumnae), a lobe-finned fish once thought extinct for 65 million years, was found alive off the coast of South Africa in 1938. Today two known populations survive in the western Indian Ocean and the waters off Sulawesi, Indonesia, representing a relict lineage that once ranged across ancient seas worldwide.
Resource Partitioning
/ REE-sors par-TIH-shun-ing / · Old French resource + Latin partitio (division)
Resource partitioning occurs when competing species reduce direct competition by using shared resources in different ways, at different times, or in different locations.
When two or more species require the same limiting resource, natural selection favors individuals that shift their use of that resource away from the zone of greatest overlap with competitors. Nile perch and tilapia in African rift lakes feed at different water depths, reducing direct dietary competition despite sharing the same body of water. Anole lizards on Caribbean islands use different perch heights and branch diameters, allowing several species to forage in the same tree without monopolizing the same microhabitat.
This division of resources permits coexistence that would otherwise end in competitive exclusion, where the superior competitor eliminates the weaker one entirely.
Robert MacArthur documented in 1958 that five warbler species in New England spruce forests all consume spruce budworms but partition the tree into distinct foraging zones, from the outermost branch tips to the inner trunk, reducing overlap enough for stable coexistence.
Competing species must always drive one another to extinction. Dividing resources along spatial, temporal, or dietary axes can reduce competitive overlap enough for multiple species to coexist indefinitely in the same habitat.
Three species of sunfish in North American lakes, the bluegill (Lepomis macrochirus), pumpkinseed (Lepomis gibbosus), and longear sunfish (Lepomis megalotis), partition foraging habitat by feeding at different water depths and on prey of different sizes. Bluegill concentrate on open-water zooplankton, while pumpkinseed specialize on hard-shelled mollusks in the littoral zone, reducing dietary overlap to less than 30 percent between the two most similar species.
Riparian Forests
/ rih-PAIR-ee-un FOR-ests / · Latin riparius (of riverbanks) + Latin forestis
Riparian forests are bands of tree and shrub vegetation growing along rivers, streams, and other watercourses, where seasonal flooding, elevated soil moisture, and proximity to flowing water create ecological conditions distinct from surrounding upland habitats.
These forests form linear corridors that connect otherwise fragmented upland habitats, allowing wildlife to move across habitats that would otherwise be impassable. Vegetation shifts from water’s edge inward through emergent aquatic plants, moisture-tolerant willows and alders, floodplain hardwoods, and transitional upland species, creating a steep gradient of plant communities within a narrow band. Cottonwood-dominated riparian corridors along the Colorado River once extended for hundreds of kilometers and supported nesting populations of southwestern willow flycatchers (Empidonax traillii extimus), a federally endangered bird that depends almost entirely on this habitat type.
Nutrient and sediment interception by riparian root systems and leaf litter can reduce nitrogen loading to adjacent streams by 50 to 90 percent compared with unforested stream banks.
Riparian forests cover less than 2 percent of land area in the western United States but support over 70 percent of vertebrate species in the region, making them the most biodiverse terrestrial habitat type per unit area in that part of the country.
Riparian forests are sometimes treated as ordinary woodland that happens to border a stream. Seasonal flooding, waterlogged anaerobic soils, and steep moisture gradients select for specialized species tolerant of physical disturbance and oxygen-poor root zones, producing plant and animal communities that differ substantially from upland forests even a short distance away.
Beavers (Castor canadensis) selectively harvest riparian willows, cottonwoods, and alders within about 100 meters of stream banks to build dams and lodges. A single beaver dam can raise the local water table enough to convert 1 to 4 hectares of dry floodplain into wetland, creating habitat used by amphibians, waterfowl, and juvenile fish.
