Ecology Terms Starting With L
Ecology Glossary: L
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Land Degradation
/ LAND deg-ruh-DAY-shun / · Old English land + Latin degradare (to lower in rank)
Land degradation is the long-term reduction in the biological productivity, ecological function, or economic usefulness of land caused by human activity or natural processes.
Degraded land loses fertile topsoil, vegetation cover, water-holding capacity, and biodiversity, often in self-reinforcing cycles that are difficult to reverse. Soil erosion removes the nutrient-rich upper horizon first; the United States loses an estimated 1.7 billion metric tons of topsoil to erosion each year, according to USDA assessments. Salinization affects roughly 20 percent of the world’s irrigated farmland, occurring when evaporation draws dissolved salts to the soil surface and concentrations reach levels toxic to most crops.
Deforestation in the Amazon Basin exposes laterite soils that harden into a brick-like layer called plinthite within a few years of canopy removal, permanently reducing agricultural potential. Restoration is possible but slow; re-establishing native grassland on degraded rangeland in the Loess Plateau of China required more than a decade of active management before soil organic matter and infiltration rates approached pre-degradation levels.
The Dust Bowl of the 1930s stripped topsoil from roughly 40 million hectares across the southern Great Plains after drought combined with deep plowing removed the native grass cover that had anchored the soil for millennia. Dust storms carried an estimated 300 million metric tons of soil eastward in a single storm on April 14, 1935, a day recorded as Black Sunday.
Land degradation only occurs in dry desert regions. Degradation affects tropical rainforests through logging and slash-and-burn agriculture, temperate farmland through nutrient depletion and compaction, and coastal wetlands through drainage and saltwater intrusion.
Desert Bird Adaptations →Overgrazing on the Sahel grasslands of sub-Saharan Africa has removed plant cover across millions of hectares since the mid-twentieth century. Without root systems to bind the soil, wind erosion advances the Sahara Desert southward at rates estimated between 48 and 100 kilometers per year in the most affected areas.
Biological Weathering 101 →Leaf Litter
/ LEEF LIT-ur / · From Old English leaf and litter meaning scattered material or bedding
Leaf litter consists of dead plant material, primarily fallen leaves, that accumulates on the soil surface and forms the base of detrital food webs in terrestrial ecosystems.
Leaf litter supports diverse communities of decomposers including fungi, bacteria, millipedes, springtails, and earthworms that break down organic matter and return nutrients to the soil. Temperate deciduous forests produce 2 to 6 metric tons of leaf litter per hectare annually, with decomposition rates varying by climate and litter chemistry. Oak leaves contain high tannin concentrations and may persist for two to three years, while sugar maple leaves decompose within one year because their carbon-to-nitrogen ratio near 50:1 supports rapid microbial activity.
The litter layer regulates soil temperature and moisture, reduces surface erosion, and provides habitat for salamanders, ground beetles, and ground-nesting birds. Nutrient release follows predictable sequences, with potassium leaching out within weeks while nitrogen and phosphorus mineralize more gradually over months to years.
A single square meter of temperate forest floor can harbor over 1,000 individual invertebrates living within the litter layer, including springtails, oribatid mites, and beetle larvae. These decomposers pass leaf fragments through their digestive systems multiple times, progressively fragmenting plant polymers and increasing the surface area available to microbial breakdown far more efficiently than microbes working alone.
Raking and removing leaf litter from forests or gardens improves soil health. Removing litter depletes soil nutrients, disrupts detrital food webs, and eliminates the moisture-retaining layer that invertebrate decomposers depend on.
Tropical rainforests of the Amazon Basin produce leaf litter year-round yet maintain litter layers rarely exceeding five centimeters in depth. Fungi and termites break down fallen leaves within six to eight weeks under the warm, humid conditions, returning nutrients to the shallow root mats that intercept them before they leach into the soil.
Lichen
/ LY-ken / · From Greek leichen meaning tree moss or what eats around itself
Lichen is a composite organism formed by a mutualistic symbiosis between a fungus and one or more photosynthetic partners, either green algae or cyanobacteria.
The fungal partner, almost always an ascomycete, provides the structural body and protects the photosynthetic cells from desiccation and ultraviolet radiation, while the algal or cyanobacterial photobionts supply carbohydrates through photosynthesis. More than 20,000 lichen species colonize extreme environments from Antarctic rocks to desert surfaces, tolerating temperatures ranging from negative 70 to positive 70 degrees Celsius. Lichens grow extraordinarily slowly; some Arctic crustose species add less than one millimeter of radius per century, a rate slow enough that scientists use lichen diameter to date glacial moraines and rock surfaces through a technique called lichenometry.
These organisms are sensitive indicators of air quality because they absorb nutrients and pollutants directly from the atmosphere, and sulfur dioxide concentrations above 30 micrograms per cubic meter eliminate the most sensitive species. Lichens contribute to primary succession by secreting organic acids that weather rock surfaces and by accumulating organic matter that eventually supports vascular plant establishment.
Lichens sent to the International Space Station in 2005 survived 15 days of unprotected exposure to space vacuum, cosmic radiation, and temperature extremes as part of the LICHENS experiment conducted by the European Space Agency. Both test species, Rhizocarpon geographicum and Xanthoria elegans, resumed normal metabolic activity after returning to Earth, suggesting that lichens could potentially survive interplanetary transport.
Lichens are plants. The fungal component comprises roughly 90 percent of the lichen body, controls reproduction, and determines the organism's form; the photosynthetic cells live embedded within fungal tissues and cannot survive independently in most environments.
Map lichen (Rhizocarpon geographicum) colonizes exposed granite boulders across Arctic and alpine regions and grows so slowly that a thallus 100 millimeters in diameter may be more than 1,000 years old. Geomorphologists use the maximum diameter of map lichen patches on glacially deposited boulders in Scandinavia and Patagonia to estimate when those surfaces were first exposed by retreating ice.
Limiting Factor
/ LIM-ih-ting FAK-ter / · Latin limitare (to set limits) + factor (maker)
Limiting Factor limiting factor is any resource or environmental condition that restricts the growth, abundance, or distribution of a population when it falls below a minimum threshold or exceeds a maximum tolerance.
Justus von Liebig articulated the core principle in 1840, observing that crop yield was constrained by whichever nutrient was in shortest supply relative to plant needs, a concept now called Liebig’s Law of the Minimum. Iron limits phytoplankton growth across large stretches of the Southern Ocean despite abundant nitrogen and phosphorus, a finding confirmed by the IRONEX experiments of the early 1990s, when adding iron to iron-poor surface water triggered phytoplankton blooms visible from satellites. Limiting factors can be biotic, such as predation pressure or pathogen load, or abiotic, such as water availability, light intensity, temperature, or dissolved oxygen.
Nest cavity availability limits populations of cavity-nesting birds like red-cockaded woodpeckers (Dryobates borealis) even in areas with abundant food, demonstrating that non-nutritional resources constrain population size just as effectively as food or water.
Ecologists distinguish between density-dependent limiting factors, whose effects intensify as population size increases, and density-independent factors, which affect populations regardless of density. Disease and competition are density-dependent; a late-season frost that kills migratory insects is density-independent. This distinction matters for predicting how populations respond to disturbance and for designing effective management strategies.
Only food limits population size. Water availability, light, temperature, nesting sites, dissolved oxygen, soil nutrients, space, predators, and disease all constrain populations, and in many ecosystems the most critical limiting factor is not food at all.
In the open ocean off the Galapagos Islands, iron concentrations in surface water drop below 0.2 nanomoles per liter, suppressing phytoplankton growth despite high macronutrient levels. During the 1993 IRONEX experiment, adding iron to a 64-square-kilometer patch of this water increased chlorophyll concentrations by a factor of 27 within ten days.
