Ecology Terms Starting With N
Ecology Glossary: N
Natal Philopatry
/ NAY-tul fih-LOP-uh-tree / · Latin natalis (of birth) + Greek philos (loving) + patra (fatherland)
Natal philopatry is returning to one's birthplace to breed. The returning animal may have traveled far in between. Natal dispersal is the opposite pattern. Those animals settle and breed away from their birthplace. Natal philopatry is common in seabirds, salmon, and sea turtles. It limits gene flow between populations.
Returning home to breed carries clear benefits. The birthplace has already proven it can raise young. Local animals also stay adapted to its conditions.
Many seabirds show colonial nesting at traditional sites. Salmon return upstream for spawning in their home river. Both depend on strong homing abilities.
Philopatry has costs as well as benefits. Relatives breeding close together face a higher risk of inbreeding. Isolated populations can also become genetically distinct.
Young Coho Salmon (Oncorhynchus kisutch) were exposed to synthetic odors. They were then released into Lake Michigan. Eighteen months later, scientists scented separate streams with each odor. Most fish returned to the stream carrying their imprinted odor.
Many assume philopatric animals never leave their birthplace. Most actually travel far before coming back. Sockeye Salmon (Oncorhynchus nerka) mature far out at sea. They typically spend 2 to 3 years there. Even so, some individuals stray and breed elsewhere.
Chinook Salmon (Oncorhynchus tshawytscha) make a very long river migration. Yukon River fish swim over 3,000 km upstream. The journey takes about three months without food. They spawn in the streams where they hatched.
What Do Salmon Eat →Nearctic
/ nee-ARK-tik / · Greek neos (new) + arktikos (northern), from arktos (bear)
Nearctic refers to the Nearctic biogeographic realm. It covers most of North America and Greenland. It includes northern Mexico's highlands and deserts. It meets the Neotropical realm in the Mexican Transition Zone. A former land bridge linked Alaska and Siberia. So it is often grouped with the Palearctic. Together they form the Holarctic.
The realm runs from Greenland and Arctic Canada into Mexico. It spans tundra, taiga, prairie, and desert. Most of the region has cold or temperate climates.
North American and Eurasian faunas share many groups. Their link was a land bridge between Siberia and Alaska. Moose and gray wolves live on both continents.
Some Nearctic animals came north from South America. Opossums, armadillos, and porcupines are famous examples. Many crossed after the Isthmus of Panama rose.
The Pronghorn (Antilocapra americana) is found only in the Nearctic. It is the last living member of family Antilocapridae. It can sprint at nearly 90 kilometers per hour.
The Nearctic and North America are not exactly the same. Tropical southern Mexico and Central America are Neotropical. The line follows animal ranges, not country borders.
The North American Porcupine (Erethizon dorsatum) is a Nearctic mammal. It ranges from central Alaska to Labrador and into Mexico. Its family, however, arose in South America. A 2-million-year-old Florida fossil, Erethizon poyeri, had a grasping tail. Living North American porcupines lack that grasping tail.
What Do Porcupines Eat →Neotropical
/ nee-oh-TROP-ih-kul / · Greek neos (new) + tropikos (of the solstice turning)
Neotropical refers to the Neotropical biogeographic realm. It extends from tropical lowland Mexico through Central America. It includes all of South America and the West Indies. Alfred Russel Wallace recognized it in 1876. He based it on shared animal distributions. Long isolation produced sloths, anteaters, and caviomorph rodents there.
The realm runs from lowland Mexico to South America’s tip. It also includes the islands of the West Indies. The drier Mexican highlands belong to the Nearctic instead.
South America drifted alone for tens of millions of years. Sloths, anteaters, and armadillos evolved there in isolation. New arrivals came by rare dispersal across open ocean.
Rodents and monkeys likely rafted over from Africa. The Isthmus of Panama later joined the two Americas. Animals then moved north and south across this land bridge.
Contamana, Peru, has rodent fossils about 41 million years old. Their ancestors likely rafted across the Atlantic from Africa. Porcupines, capybaras, and guinea pigs descend from these colonists.
Neotropical does not simply mean hot and tropical. The realm is defined by shared animal groups, not climate. It even reaches the cold subantarctic zone of South America.
The Brazilian Porcupine (Coendou prehensilis) is a Neotropical mammal. Its range runs from eastern Venezuela to northern Argentina. It also lives on the island of Trinidad. It occurs from lowlands up to about 1,500 meters.
What Do Porcupines Eat →Nest Predation
/ NEST preh-DAY-shun / · Old English nest + Latin praedatio (plundering)
Nest predation is the loss of nest contents to predators. Predators may take eggs, nestlings, or sometimes a brooding parent. Snakes, mammals, and other birds are common nest predators. It is the leading cause of songbird nest failure. Biologists often report it as a daily nest loss rate.
Nest predation shapes how songbirds evolve their life histories. Many birds hide nests in dense vegetation. Others nest in cavities or high above the ground.
Where a nest sits matters greatly. Poor nest sites tend to be found and raided quickly. Experiments show these site effects repeat from year to year.
Human activity can add new nest predators. Introduced predators threaten birds that evolved without them. American mink and feral cats now roam Chile’s Navarino Island.
On Chile’s Navarino Island, camera traps watched songbird nests. The Chimango Caracara (Milvago chimango) was the top nest predator. It raided 13 of 15 nests with an identified predator. Predation caused 71 percent of nest failures there.
Many assume a nest is safest once the chicks hatch. That is not always true. Missouri Indigo Buntings (Passerina cyanea) lost more nests after hatching. On Navarino Island, 69 percent of raids hit nestlings.
Missouri Field Sparrows (Spizella pusilla) were studied for 7 years. Researchers tracked nests in old fields alongside indigo buntings. Nest height was the only habitat feature that consistently mattered. Nests over 3 meters high almost always fledged young.
Branches of Ecology →Niche
/ NITCH / · French niche (recess in a wall)
Niche is the full set of biotic and abiotic conditions a species requires to survive and reproduce, including the resources it uses, the environmental tolerances it has, and its interactions with other organisms.
A species occupies a fundamental niche defined by all physical and chemical conditions it can tolerate, such as a temperature range of 5 to 35 degrees Celsius and a pH range of 6 to 8. Competition from other species, predation pressure, and parasitism typically restrict where the species actually survives, producing a realized niche that is smaller than the fundamental niche. G.
Evelyn Hutchinson formalized this distinction in 1957, describing the niche as an n-dimensional hypervolume in which each axis represents a different environmental variable. A pileated woodpecker’s (Dryocopus pileatus) niche includes foraging for wood-boring insects under loose bark, roosting in dead snags, and excavating nest cavities in dead wood, activities that shape forest structure for many other species.
The concept of competitive exclusion, formalized by Georgy Gause in 1934 using laboratory cultures of Paramecium, predicts that two species with identical niches cannot coexist indefinitely. One species will always outcompete the other, driving it locally extinct or forcing a shift in resource use.
Niche means only where an animal lives. Habitat is place, while niche is the species' role and requirements in that place.
The red-cockaded woodpecker (Leuconotopicus borealis) occupies a highly specialized niche in longleaf pine (Pinus palustris) forests of the southeastern United States, excavating nest cavities exclusively in living pines infected with red heart fungus. Colonies require territories of 80 to 200 hectares of mature pine forest, and their cavity trees are used by at least 27 other vertebrate species after the woodpeckers abandon them.
What Do Woodpeckers Eat? →Niche Overlap
/ NITCH OH-ver-lap / · French niche (recess) + Old English oferhleopan (to overlap)
Niche overlap is the degree to which two or more species share the same resources, environmental conditions, or habitat space within an ecosystem.
When two species exploit the same food types, feeding locations, or microhabitats, their niches overlap and interspecific competition intensifies as shared resources become limited. Robert MacArthur’s classic 1958 study of five warbler species in northeastern spruce forests showed that each species foraged in a distinct vertical zone within the same trees, reducing overlap and allowing coexistence. Coexistence despite niche overlap is possible when resource abundance exceeds total demand, when species differ along other niche dimensions, or when they partition resources temporally by feeding at different times of day.
Measuring niche overlap quantitatively, ecologists use indices such as Pianka’s index, which ranges from 0 (no overlap) to 1 (complete overlap).
Niche overlap and competitive exclusion are linked by the limiting similarity concept, which proposes that species can coexist only if their niches differ by at least a minimum amount. Theoretical and empirical work suggests that coexisting competitors typically differ by a factor of roughly 1.3 in the size of food items they consume, a ratio sometimes called Hutchinson's ratio.
Species with niche overlap cannot coexist in the same location. Overlapping species can coexist if resources are abundant, if they differ in other niche dimensions, or if they partition resources through space or time.
African lions (Panthera leo) and spotted hyenas (Crocuta crocuta) on the Serengeti overlap substantially in diet, both targeting wildebeest and zebra. Despite this overlap, stable coexistence persists because lions kill prey averaging about 230 kilograms while hyenas more frequently take prey in the 50 to 150 kilogram range, and the two species also differ in activity timing, with hyenas doing most hunting after dark.
Nitrogen Cycle
/ NY-troh-jen SY-kul / · Latin nitrum (niter) + Greek kyklos (circle)
Nitrogen Cycle is the biogeochemical pathway by which nitrogen moves and changes form among the atmosphere, soil, water, and living organisms.
Nitrogen fixation by bacteria such as Rhizobium and free-living Azotobacter converts atmospheric nitrogen gas into ammonium, making nitrogen biologically available for the first time in each cycle. Nitrification by Nitrosomonas and Nitrobacter bacteria then converts ammonium to nitrite and nitrate in soil and water, the forms most plants absorb through their roots. Animals obtain nitrogen by consuming plants or other organisms, and decomposers break down organic nitrogen in dead tissue back into ammonium through ammonification.
Denitrification by bacteria such as Pseudomonas returns nitrogen gas to the atmosphere, completing the cycle; globally, biological nitrogen fixation adds roughly 120 million metric tons of nitrogen to terrestrial ecosystems each year.
Industrial nitrogen fixation through the Haber-Bosch process, developed by Fritz Haber and Carl Bosch in the early twentieth century, now fixes more nitrogen annually than all natural biological fixation combined. This synthetic nitrogen feeds roughly half the world's human population through increased crop yields but has also driven widespread nitrogen pollution in rivers, coastal waters, and the atmosphere.
Plants use nitrogen gas directly from air the way they use carbon dioxide. Most plants require nitrogen in reduced forms such as ammonium or nitrate that bacteria or synthetic fertilizers provide.
Alder trees (Alnus species) form mutualistic root nodules with the nitrogen-fixing bacterium Frankia, allowing them to colonize nitrogen-poor soils such as glacial outwash and landslide scars. A single alder stand can add 50 to 100 kilograms of fixed nitrogen per hectare per year to the surrounding soil, accelerating forest succession by enriching conditions for other plant species.
Not Evaluated
/ NOT ih-VAL-yoo-ay-tid / · English not + evaluate, from French évaluer, from Latin valere (to be worth)
Not Evaluated is an IUCN Red List category. It applies to taxa never assessed against the criteria. It says nothing about extinction risk. A Not Evaluated species may be safe or threatened. Most described species on Earth fall into this category.
Every newly described species begins without a Red List assessment. It keeps this label until assessors formally review it. Reviews then apply the same five criteria, A to E.
By 2025, the Red List included 172,620 species. That is still a small share of described species. Birds and mammals are far better covered than insects.
Taxonomic splits can instantly create unassessed species. Unassessed species can be seriously threatened without anyone knowing. IUCN warns against treating them as not threatened.
IUCN had assessed only about 8 percent of described species. The 2025 figure for insects was about 1 percent. Every known bird species, by contrast, has been assessed.
Not Evaluated is often confused with Data Deficient. A Data Deficient species was assessed but lacked data. A Not Evaluated species has had no assessment at all.
The Amazonian Long-tailed Porcupine (Coendou longicaudatus) is Not Evaluated. A 2021 study split it from the Brazilian Porcupine. It ranges across much of South America. No Red List assessment covers it yet.
What Do Porcupines Eat →Nutrient Cycling
/ NYOO-tree-ent SY-kling / · Latin nutrire (to nourish) + Greek kyklos (circle)
Nutrient cycling is the repeated movement of chemical elements such as carbon, nitrogen, phosphorus, and sulfur between living organisms and the nonliving environment through biogeochemical processes.
Decomposer organisms including bacteria, fungi, and soil invertebrates break down dead organic matter and release nutrients back into soil and water, where plant roots and microbes can reabsorb them. This recycling distinguishes nutrient dynamics from energy flow: matter cycles indefinitely through ecosystems, while energy enters as sunlight and exits as heat and cannot be reused. Phosphorus cycling illustrates how nutrients can become temporarily unavailable; phosphorus locked in deep ocean sediments or in mined rock takes millions of years to return to the surface through geological uplift, making it a limiting nutrient in many freshwater and terrestrial systems.
Mycorrhizal fungi accelerate nutrient cycling by extending the absorptive surface of plant roots and releasing enzymes that break down organic phosphorus and nitrogen compounds in soil.
Tropical rainforests cycle nutrients so rapidly that most of the ecosystem's nutrient capital resides in living biomass rather than in soil. When rainforest is cleared and burned, the nutrient pulse released by ash is exhausted within a few growing seasons, leaving soils that are often too poor to sustain agriculture without continuous fertilizer inputs.
Nutrients disappear after organisms use them and must be replaced. Chemical nutrients cycle repeatedly through living and nonliving ecosystem components, though they can become temporarily unavailable in certain compartments.
In a temperate deciduous forest, a single hectare of soil can contain more than a metric ton of fungal mycelium actively decomposing leaf litter and woody debris. White rot fungi such as Trametes versicolor break down lignin in fallen logs, releasing carbon, nitrogen, and phosphorus that tree roots and soil bacteria absorb within the same growing season.
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